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Marine Biology (Pechenik textbook)
Flashcards to accompany Jan Pechenik's "Biology of the Invertebrates"
188
Biology
Undergraduate 3
01/26/2013

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Term
Ammonia
Definition
The basic end product of amino acid metabolism in all organisms, regardless of habitat. Ammonia is usually very toxic, largely through its effects on cellular respiration.
Term
Phytoplankton
Definition
Single-celled organisms (plants)
Term
Zooplankton
Definition
single-celled organisms (animals)
Term
suspension-feeding
Definition
Feeding without moving - suspension feeders capture food particles directly from the liquid medium that surrounds them as the particles flow past the stationary animal
Term
primary production
Definition
fixation of carbon from carbon dioxide into carbohydrates
Term
sessile
Definition
non-moving
Term
viscosity
Definition
essentially measures the extent to which the molecules of a fluid stick to one another
Term
Reynolds numbers
Definition
essentially a ratio of inertial to viscous forces
Term
drag
Definition
frictional resistance
Term
hyperosmotic
Definition
the state of being higher in osmotic concentration than one's surroundings
Term
osmotic equilibrium
Definition
when the concentration of solutes in animals' body fluids are equivalent to the concentration of solutes in the fluids that surround them
Term
metazoans
Definition
the oldest known fossils of multicellular animals. True animals are multicellular, generally diploid organisms and each develops from a blastula. They are also referred to as the Metazoa.
Term
Cambrian explosion
Definition
the amazingly sudden appearance and apparently rapid diversification of complex animals over several millions of years (around 543 million years ago)
Term
blastula
Definition
An animal embryo at the early stage of development. Also called blastosphere.
Term
unicellular
Definition
single-celled
Term
acellular
Definition
without cells
Term
bilaterally symmetrical
Definition
possessing right and left sides that are approximately mirror images of one another
Term
cephalization
Definition
the concentration of nervous and sensory tissues and organs at one end of an animal, resulting in distinct anterior and posterior ends
Term
radial symmetry
Definition
Radially symmetrical organisms can be divided into two approximately equal halves by any cut that passes through their center
Term
Germ layers
Definition
groups of cells that behave as a unit during the early stages of embryonic development and give rise to distinctly different tissue and/or organ systems in the adult
Term
diploblastic animals
Definition
these animals have only two distinct germ layers that form during or following the movement of cells into the embryo's interior
Term
ectoderm
Definition
the outermost layer of cells in a diploblastic animal during the embryonic phase
Term
endoderm
Definition
the innermost layer of cells in a diploblastic animal during the embryonic phase
Term
triploblastic
Definition
Most metazoans are triploblastic. During the ontogeny of triploblastic animals, cells of either the ectoderm or, more usually, the endoderm give rise to a third germ layer, the mesoderm.
Term
mesoderm
Definition
The mesodermal layer of tissue always lies between the outer ectodermal tissue and the inner endodermal tissue.
Term
acoelomate
Definition
Triploblastic animals that lack an internal body cavity
Term
blastocoel
Definition
an internal space that develops in teh embryo prior to gastrulation. This type of body cavity is termed a pseudocoel, and the organism housing it is said to be a pseudocoelomate.
Term
psdeudocoel
Definition
a body cavity like a pseudocoel
Term
pseudocoelomate
Definition
an animal that possesses a pseudocoel
Term
coelom
Definition
an internal, fluid-filled body cavity lying between the gut and the outer body wall musculature and lined with tissue derived from embryonic mesoderm
Term
coelomates
Definition
animals that possess coeloms. They are also known as eucoelomates.
Term
protostomes
Definition
In these animals, coelom formation occurs by gradual enlargement of a split in the mesoderm. This process is termed schizocoely.
Term
schizocoely
Definition
coelom formation that occurs by gradual enlargement of a split in the mesoderm.
Term
deuterostomes
Definition
In these animals, the coelom typically forms through envagination of the archenteron into the embryonic blastocoel. Because the coelom of deuterostomes forms from a part of what eventually becomes the gut, coelom formation in this group of animals is termed enterocoely.
Term
enterocoely
Definition
In these animals that show enterocoely, the coelom typically forms through envagination of the archenteron into the embryonic blastocoel and forms part of what eventually becomes the gut.
Term
animal pole
Definition
Generally, yolk is asymmetrically distributed within eggs, and the nucleus occurs in, or moves to, the region of lower yolk density. This is the animal pole, and it is here that the polar bodies are given off during meiosis.
Term
vegetal pole
Definition
The opposite end of the egg from the animal pole. Generally, yolk is asymmetrically distributed within eggs, and the nucleus occurs in, or moves to, the region of lower yolk density. This is the animal pole, and it is here that the polar bodies are given off during meiosis.
Term
radial cleavage
Definition
Happens in deuterostomes. The spindles of a given cell, and thus the cleavage planes, are oriented either parallel or perpendicular to the animal-vegetal axis. Thus, daughter cells derived from a division in which the cleavage plane is parallel to the animal-vegetal axis end up lying in the same plane as the original mother cell.
Term
spiral cleavage
Definition
The spindle axes of cells undergoing spiral cleavage are oriented (after the first two cleavages) at 45 degree angles to the animal-vegetal axis. Moreover, the division line does not necessarily pass through the center of the dividing cell.
Term
micromeres
Definition
Smaller cells that occur at the eight-cell stage of spiral cleavage and lie in the spaces between the underlying larger cells.
Term
macromeres
Definition
larger cells that occur during the 8-cell stage of cleavage
Term
indeterminate (or regulative) cleavage
Definition
Deuterostomes are said to show indeterminate/regulative cleavage -- each cells retains -- sometimes as late as the eight-cell stage -- the capacity to differentiate teh entire organism if that cell loses contact with its associatives.
Term
determinate/mosaic cleavage
Definition
Cleavage shown by protostomes. The development potential of each cell is irrevocably determined at the first cleavage; separate the blatomeres of a two-celled protostome embryo and each cell will, in most species, give rise only to a "short-lived, malformed monster."
Term
polar lobes
Definition
A conspicuous bulge of cytoplasm that forms prior to cell division and that contains no nuclear material.
Term
taxon
Definition
Any named group of organisms (eg sea urchins, banana slugs) that is sufficiently distinct to be assigned to such a category.
Term
monophyletic
Definition
At every taxonomic level where all the members of any particular phylum are presumed to have evolved from a single ancestral form. Most modern workers now agree that all monophyletic groups also must include all descendants of the originating ancestor.
Term
paraphyletic
Definition
A group that does not adhere to the following stipulations: At every taxonomic level where all the members of any particular phylum are presumed to have evolved from a single ancestral form. Most modern workers now agree that all monophyletic groups also must include all descendants of the originating ancestor.
Term
species
Definition
Theoretically, the members of one species are reproductively isolated from members of all other species.
Term
scientific name of a species
Definition
The scientific name of a species is binomial (has two parts): the generic name and the specific name. The generic and specific names (ie the species name) are usualy italicized in print and underlined in writing. The generic name begins with a capital letter, but the specific name does not.
Term
PhyloCode
Definition
Unlike the Linnaean system, the PhyloCode -- although it remains hierarchical -- is rankless: there will be no classes, orders, or families.
Term
convergence
Definition
The process through which distantly related animals may come to resemble each other rather closely.
Term
analogous
Definition
Features that resemble each other through convergence are referred to as analogous, as opposed to homologous
Term
Homologous
Definition
Morphological features that share a common evolutionary origin are said to be homologous
Term
polarity
Definition
The direction of evolutionary change
Term
ancestral state
Definition
The original state of of a character (trait?)
Term
derived state
Definition
The more advanced state of a character (trait?)
Term
clade
Definition
A group of organisms that includes the most recent common ancestor of all its members and all descendants of that ancestor; every valid clade forms a monophyletic group
Term
cladogenesis
Definition
the splitting of a single lineage into 2 or more distinct lineages
Term
anagenesis
Definition
change occuring within a lineage
Term
cladogram
Definition
the pictoral representation of branching sequences that are characterized by particular changes in key morphological or molecular characteristics (character states)
Term
homologous characters, homology
Definition
Characters that have the same evolutionary origin from a common ancestor, often coded for by the same genes. Homology is the basis for all decisions about evolutionary relationships among species
Term
taxon
Definition
any named group of organisms, such as jellyfish or sea urchins
Term
monophyletic taxon
Definition
a group of species that evolved from a single ancestor and includes all descendants of that ancestor. By definition, every valid clade must form a monophyletic taxon.
Term
parsimony
Definition
a principle stating that, in the absence of other evidence, one should always accept the least complex scenario
Term
polarity
Definition
the direction of evolutionary change
Term
ancestral/primitive state
Definition
the character state exhibited by the ancestor from which current members of a clade have evolved. Also called the plesiomorphic state
Term
derived state
Definition
an altered state, modified from the original or ancestral condition. Also called the apomorphic state
Term
apomorphy
Definition
any derived or specialized character
Term
pleisiomorphy
Definition
any ancestral or primitive character
Term
synapomorphy
Definition
a derived character taht is shared by the most recent common ancestor and by 2 or more descendants of that ancestor. In cladistic methodology, synapomorphies define clades; that is, they determine which species or other groups are most closely related to one another. Essentially, synapomorphies are homologous characters that define clades.
Term
autapomorphy
Definition
A derived character possessed by only one descendant of an ancestor, and thus of no use in discerning relationships among other descendants
Term
homoplasy
Definition
the independent acquisition of similar characteristics (character states) from different ancestors through convergence or parallelism. Such homoplastic events create the illusion of homology.
Term
paraphyletic grouping
Definition
a group of species sharing an immediate ancestor but not including all descendants of that ancestor.
Term
polyphyletic grouping
Definition
an incorrect grouping containing species that descended from 2 or more different ancestors. Members of polyphyletic groups do not all share the same immediate ancestor. Members of polyphyletic groups may resemble each other because of the independent evolution of similar traits by different ancestors.
Term
bootstrapping
Definition
a technique for evaluating the reliability of a branch of a phylogenetic tree by resampling some number of characters from teh original data set (with replacement) at random.
Term
Bayesian inference
Definition
a statistical technique used to infer the probability that a particular phylogenetic hypothesis is correct
Term
jackknifing
Definition
A technique for evaluating the reliability of a branch of a phylogenetic tree by deleting some percentage of information (eg base-pair position information) at random and then rerunning the analysis.
Term
node
Definition
a branching point on a cladogram
Term
synapomorphies
Definition
shared characteristics derived from a common ancestor in which the characters originated
Term
outgroup
Definition
the direction of evolutionary change, polarity, is first determined by comparison with a closely related taxon, the outgroup, that lies outside the taxa being studied
Term
sister groups
Definition
groups derived from the same ancestor
Term
antagonize
Definition
to act against
Term
hydrostatic skeleton
Definition
a skeleton in which fluid serves as the vehicle through which sets of muscles interact
Term
longitudinal muscles
Definition
muscles that run lengthwise down the body of an animal
Term
circular muscles
Definition
muscles that circle the body of an animal
Term
setae/chaetae
Definition
chitinous/calcareous bristles
Term
vermiform
Definition
worm-shaped
Term
metamerism/metameric segmentation
Definition
A serial repetition of segments and organ systems (skin, musculature, nervous system, circulatory system, reproductive system, and excretory system)
Term
septa
Definition
thin sheets of mesodermally derived tissue
Term
peritoneum
Definition
mesodermally derived tissue
Term
nephridia
Definition
A tubule open to the exterior that acts as an organ of excretion or osmoregulation.
Term
metanephridium
Definition
Nephridia that are open at both ends
Term
nephrostome
Definition
Nephrostome is the funnel-like component of a metanephridium. It is always oriented towards the coelom.
Term
tentacles
Definition
sensory appendages
Term
prostomium
Definition
the anterior-most part of the body of an annelid
Term
parapodia
Definition
thin, flattened outgrowths; One of the fleshy paired appendages of polychete annelids that function in locomotion and breathing.
Term
acicula
Definition
a needle-like bristle that provides internal support for the appendages (chaetae) of some polychaete worms
Term
elytra
Definition
overlapping protective plates that cover the bodies of some species
Term
perforate septa
Definition
Septa that are between anterior adjacent segments in annelids that are absent or incomplete
Term
proboscis
Definition
a polychaete's everted pharynxe
Term
errant species
Definition
a generally active, mobile species
Term
protonephridia
Definition
A protonephridium ('proto' = first) is a network of dead-end tubules lacking internal openings; pressurization drives waste fluids from the inside of the animal, and they are pulled through small perforations in the terminal cells and into the protonephridium.
Term
epitoky
Definition
a marked morphological transformation in preparation for reproductive activity
Term
epitoke
Definition
a sexually mature being, male or female, that is highly specialized for swimming and sexual reproduction
Term
atoke
Definition
during epitoky, one or more new, reproductive modules, epitokes, are budded, one segment at a time, from the posterior portion of the original animal, the atoke
Term
prototroch
Definition
a preoral circlet of cilia of a trochophore larva.
Term
telotroch
Definition
the preanal tuft of cilia in a trochophore larva.
Term
pygidium
Definition
The terminal part or hind segment of the body in certain invertebrates.
Term
trochophore
Definition
The planktonic larva of certain invertebrates, including some mollusks and polychaete worms, having a roughly spherical body, a band of cilia, and a spinning motion. NOTE polychaetes have two rings, and a third ring that forms later between the prototroch and the telotroch
Term
ramet
Definition
branch
Term
hermaphroditic
Definition
both male and female reproductive apparatus are contained within a single individual
Term
clitellum
Definition
a specialized region of the epidermis; A raised band encircling the body of oligochaete worms and some leeches, made up of reproductive segments.
Term
prostromium
Definition
the anteriormost region of the body
Term
peristaltic waves
Definition
a series of localized contractions and relaxations of the circular and longitudinal musculature
Term
retrograde waves
Definition
peristaltic waves that travel in the opposite direction of an animal's movement
Term
direct waves
Definition
peristaltic waves that travel in the same direction as an animal's movement
Term
spermathecae
Definition
specialized organs that store sperm
Term
parthenogenetic
Definition
the state of having eggs that may develop normally in the absence of fertilization
Term
mesenchyme
Definition
connective tissue that can fill a continuous coelomic space
Term
gametogenesis
Definition
gamete production
Term
spermatophores
Definition
packets of sperm
Term
clitellum
Definition
generally functions in producing the cocoon and a nutritive fluid, as in the oligochaetes
Term
digestive glands/ digestive caeca
Definition
some species form them with evaginations of the gut
Term
crop
Definition
used for food storage within body
Term
gizzard
Definition
a highly muscular structure lined with hardened cuticle, for grinding food
Term
Calciferous glands
Definition
associated with the esophagus; these may function in regulating blood pH by controlling hte concentration of carbonate ions
Term
typhlosole
Definition
the intestine of many terrestrial oligochaete species is thrown into a ridge or fold, typhlosole, which increases the gut's effective surface area
Term
chloragogen
Definition
associated with the intestine and dorsal blood vessels of oligochaetes; it is a characteristic yellow tissue; chloragogen cells play major roles in protein, carbohydrate, and lipid metabolism of oligochaetes
Term
hirudin
Definition
an anticoagulant secreted by leeches' salivary glands
Term
hermaphroditic
Definition
both male and female reproductive apparatus are contained within a single individual
Term
clitellum
Definition
the clitellum secretes a cocoon within which embryos develop. It also secretes a mucus taht assists in transferring sperm between individuals and produces albumen
Term
albumen
Definition
serves as a food source for embryos as they develop within the cocoon of a clitella species
Term
metazoans
Definition
the oldest known fossils of multicellular animals; about 543-635 million years old
Term
PCR
Definition
Polymerase Chain Reaction. Developed in mid-1980s. Permits biologists to very quickly and inexpensively generate many copies of specific DNA sequences
Term
homology
Definition
morphological features that share a common evolutionary origin
Term
polarity
Definition
the direction of evolutionary change
Term
original state
Definition
the ancestral state of a feature
Term
advanced state
Definition
the derived state of a feature
Term
tagmatization
Definition
specialization of groups of segments in animals
Term
epicuticle
Definition
the outermost layer of an arthropod; it is generally waxy, being composed of a firm lipoprotein layer underlain by layers of lipid
Term
proticule
Definition
composes teh bulk of the exoskeleton; it is composed of the polysaccharide chitin in association with a number of proteins
Term
sclerotization
Definition
Hardening is accomplished by "tanning" the proticule's protein component. The tanning process, also called sclerotization, involves the cross-linkages between protein chains; this contributes to cuticle hardening in all arthropods
Term
hemocoel
Definition
The primary body cavity of most invertebrates, containing circulatory fluid; the hemocoel also acts as an internal hydrostatic skeleton until the new exoskeleton hardens
Term
ecdysis
Definition
The process of removing the existing exoskeleton
Term
biomass
Definition
The total quantity or weight of organisms in a given area or volume
Term
Y-organ
Definition
a gland located in the head of crustaceans
Term
prothoracic glands
Definition
glands located in the thorax of insects
Term
ecdysteroid hormones
Definition
hormones that stimulate molting
Term
x-organ
Definition
a neurosecretory complex located in the eyestalks that inhibits ecdysteroid production
Term
chromatophore cells
Definition
a pigment-bearing cell; especially : one of the cells of an animal integument capable of causing integumentary color changes by expanding or contracting
Term
ostia
Definition
perforations in the heart wall that allow blood to enter the heart directly from the hemocoel
Term
ocellus
Definition
a small cup with a light-sensitive surface backed by light-absorbing pigment
Term
ommatidia
Definition
the compound eye is composed of many individual units called ommatidia
Term
crystalline cone
Definition
serves as a lens in some insects and in most crustaceans
Term
retinular cells
Definition
contain light-sensitive pigment; they are cylindrical
Term
shielding pigment
Definition
potically isolates every ommatidium from surrounding ommatidia
Term
neural cartridge
Definition
a cluster of neurons receiving the information carried by the retinular cells and sending action potentials to the optic ganglia for processing
Term
rhabdomeres
Definition
find, microvillar outfoldings of the retinular cell walls
Term
rhabdom
Definition
The rhabdomeres within each ommatidium form a discrete, ordered association called a rhabdom
Term
apposition eye
Definition
A type of compound eye occurring in diurnal insects, in which each ommatidium is surrounded by a shield of pigment; the lens is directly apposed to the receiving rhabdom
Term
superposition eye
Definition
In such an eye, each ommatidium has a large space between the distal end of the crystalline cone and the rhabdom; without the shielding pigment in the way, light from a single point in the visual field can be received by many lenses and focused into a single rhabdom, producing a signal of significantly greater intensity than received through a single lens
Term
thorax
Definition
The part of the body of a mammal between the neck and the abdomen, including the cavity enclosed by the ribs, breastbone, and dorsal...
The corresponding part of a bird, reptile, amphibian, or fish.
Term
carapace
Definition
a bony or chitinous case or shield covering the back or part of the back of an animal (as a turtle or crab)
Term
cephalothorax
Definition
The fused head and thorax of spiders and other chelicerate arthropods
Term
rostrum
Definition
in some species, the carapace bears a prominent anterior projection called the rostrum
Term
compound eyes
Definition
An eye consisting of an array of numerous small visual units, as found in insects and crustaceans.
Term
mandible
Definition
The jaw or a jawbone, esp. the lower jawbone; they crush food
Term
maxillae
Definition
part of the mouth; generates water currents and manipulates food
Term
maxillipeds
Definition
(in crustaceans) An appendage modified for feeding, situated in pairs behind the maxillae.
Term
walking legs
Definition
commonly known as pereopeds
Term
chelate
Definition
claw-bearing
Term
pleopods
Definition
the first 5 pairs of abdominal appendages
Term
uropods
Definition
the last pair of abdominal appendages
Term
biramous
Definition
possessing two branches
Term
endopodite
Definition
the inner branch of a malacostracan appendage
Term
exopodite
Definition
the outer branch of a malacostracan appendage
Term
chromatophores
Definition
highly branched cells containing pigment granules
Term
sinus gland
Definition
hormones are manufactured by the so-called X-organ located in the eyestalks and are transported a short distance to the sinus gland for storage
Term
maxillipeds
Definition
An appendage modified for feeding, situated in pairs behind the maxillae.
Term
photophores
Definition
light-producing organs
Term
carina, rostrum, scuta, terga
Definition
different plates that compose the shells of barnacles
Term
nauplius
Definition
The first larval stage of many crustaceans, having an unsegmented body and a single eye; typical of several diverse groups of crustaceans, including the copepods, ostracods, branchiopods, euphausiids and cirripedes
Term
copepodite
Definition
“The nauplius undergoes five moults before it transforms into a copepodite stage that more closely resembles the adult.”
Term
cypris stage
Definition
the final stage of barnacle larval development following the planktonic nauplius stage. the cypris settles on a hard substrate and undergoes metamorphosis into benthic adult form. The larva is also known as a cyprid.
Term
kentrogon
Definition
a larva of a parasitic barnacle (order Rhizocephala); the kentrogon is specialized for piercing through the host cuticle
Term
phytoplankton
Definition
unicellular, free-floating, photosynthesizing protists
Term
zooplankton
Definition
Plankton consisting of small animals and the immature stages of larger animals.
Term
meiobenthos
Definition
the community of small animals living in association with sediment
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