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Weed Science: Midterm 1
University of Guelph CROP*4240
274
Agriculture
Undergraduate 4
09/28/2017

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Cards

Term
Aerial bulblet
Definition

Bulbil

An organ used for vegetative propagation by some creeping perennials. Includes wild garlic. Bulbils are produced in the inflorescence, and can be carried much further from the plant than bulbs.

Term
Agriculture
Definition
It goes against the natural course of nature. The land has a high level of disturbance, affecting of all its existing vegetation. Fertilizers are added, so there are more nutrients than in the natural environment; this can cause competition.
Term
Allelopathy
Definition

Direct interference

A contributing factor of interference. The negative effect that one plant exerts on another by producing chemical substances. Allelopathic toxins can be released by: exudation from roots, washing from leaves, volatile gases, and decaying plant parts. It is difficult to prove whether a plant is allelopathic or not, because we don't know all the weeds that use allelopathic compounds. Some aggressive perennials release toxic compounds from decaying dead material, which can be a problem if susceptible crops are planted on a site that was previously heavily infested by these weeds. Evolved to help plants reduce competition for resources. Gives advantage to some weeds.

Term
Allergies and toxicities
Definition
A negative aspect of weeds. Can affect humans and animals. Includes skin irritation from poison ivy, giant hogweed, and wild parsnip. Many people have allergies to pollen from ragweed.
Term
Amaranth
Definition
Family Amaranthaceae. An edible plant; the seeds and leaves are edible. High in fibre, and gluten free.[image]
Term
Amaranthaceae
Definition
A family of plants. The pigweed and amaranth family. Includes redroot pigweed, green pigweed, and amaranth.
Term
Animals
Definition
A form of seed dispersal. Some seeds can stick to the fur of animals, and may stick to livestock, pets, and people's clothing. They can get into the ears, eyes, and mouths of animals, and may cause discomfort.
Term
Annual
Definition
A plant which completes its lifecycle in one year or less. Includes cool season and warm season annuals. Can respond to sudden fluxes in nutrients, such as after a fire, or in agricultural fields, by accumulating nitrogen quickly. Cannot compete later on with perennial plants.
Term
Aphids
Definition
The action threshold is 250 insects per plant, at V4 in soybeans.
Term
Apical dominance
Definition
The growing point of a plant produces the hormone auxin (IAA), which moves down the organ, preventing formation of lateral branches. Lateral buds are "dominated" by the apex, and stay dormant. Ensures direction in growth. If you break the tip of the organ, it prevents auxin from reaching the buds, and they can now grow and produce new shoots. This method can be used to make ornamental plants bushier. Helps perennials recover from damage, such as from fire.
Term
Apomixis
Definition

Agamospermy

Production of seeds without fertilization. A form of asexual reproduction. There are several types. The flower may require contact with pollen, but there is no exchange of gametes. The seed has the same genome as the mother plant; it is a clone. Some plants are evolved to have apomixis more often than others. Occurs in cypress, citrus crops, and many arctic species. Can cause problems for plant breeders.

Term
Aquatic weeds
Definition
Weeds which grow in water. Can block channels and lakes, causing problems with boating and swimming activities. Can appear in garden ponds. Tend not to interfere with agriculture, except in flooded crops such as rice and cranberries. May be floating or submerged. There are herbicides for aquatic weeds which are poured into the water.
Term
Arabidopsis
Definition

Family Brassicaceae. When grown under low red/far red light ratio, similar to shade, it has a narrow growth habit compared to a plant grown under normal light conditions, even when light intensity is the same.

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Term
Artificial dispersl
Definition
Seed dispersal helped by human activities. Includes contaminated seed (crop seed not properly cleaned), machinery, contaminated feed, and contaminated soil or potting mix. Seeds can travel in tiny nooks inside machinery, especially combines, or on mud stuck to wheels. Ideally, combines should be cleaned after each use, but this is unrealistic. Grain is harder to clean when weed seeds are a similar size or shape to the crop seeds.
Term
Asexual reproduction
Definition

Clonal reproduction

Includes apomixis and vegetative reproduction (rhizomes, plagiotropic roots, stolons, tubers, bulbs). The plants benefit from genetic stability of clones: plants with best adaptations to the environment remain in place. A simpler way to reproduce that does not require the same energy investment as sexual reproduction, such as elaborate flowers and specialized organs. Represents an additional way to reproduce that may supplement seed production. May allow some species to reproduce and multiply under sub-optimal conditions. Some species may have both sexual and asexual reproduction under optimal conditions, but at the margin of its natural distribution range, where conditions are sub-optimal, it has only asexual reproduction. Even if sexual reproduction is impaired, asexual reproduction may still take place. A disadvantage is that there is no exchange of genetic material. Does not increase genetic diversity, the basis for natural selection. A major environmental change, such as herbicide application, may negatively affect the whole population.

Term
Autogenic
Definition
"Within"
Term
Banana
Definition

Musa sp.

Family Musaceae. Has vegetative propagation with ratoons.

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Term
Beggarticks
Definition

Bidens sp.

Family Asteraceae. The seeds have little appendages which look like legs, making them resemble ticks.

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Term
Benefits of weeds
Definition
Many species can be a food source for wildlife, and most weeds have high nutritional value. In some circumstances, they can serve as emergency feed for cattle. Some have very vigorous underground growth and can help reduce soil erosion. In addition to high biomass production, they can increase soil organic matter. Many species have been investigated as potential food or base material for industry. Can be a source of genetic material for plant breeding, providing genes for resistance to diseases or environmental stress. Can serve as a refuge for beneficial insects, or as a trap for pests. Some have good aesthetic qualities. Have nutrient retention and can prevent of leaching. Can shade horticultural crops. Have many of the good qualities of cover crops. May be edible or have medicinal uses. Increase biodiversity.
Term
Biennial
Definition
A plant which takes two years to finish its lifecycle. The first year it builds up biomass, and the second year it flowers. This definition may be challenged by some botanists because biennials' lifecycles can be variable, up to four years.
Term
Biotechnology
Definition
In the 1980s when biotechnology was still new, people thought that we would be able to produce perennial wheat, or nitrogen-fixing grasses, but it didn't happen.
Term
Birds
Definition
Some seeds are dispersed by being eaten by birds. The digestive tract helps wear down the seed coat enough for germination to occur. When the bird poops the seeds out, it disperses them.
Term
Black medick
Definition

Medicago lupulina

Family Fabaceae. A legume. The first true leaf is an oval unifoliate. Later leaves are trifoliate, with diamond-shaped leaflets. The centre leaf has a petiole whereas the other two do not. Has a prostrate growth habit. Found in turf and pastures. Has good qualities as feed for livestock. Adapted to less disturbed environments. Has clusters of small yellow flowers. The name comes from its black seed pods. Can look similar to common yellow woodsorrel.

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Term
Broad-leaved plantain
Definition

Plantago major

Family Plantaginaceae. A stationary perennial. Has vegetative propagation with a shallow underground shoot, and ratoons. Has a short vertical stem.

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Term
Buckthorn
Definition

Rhamnus sp.

Family Rhamnaceae. An invasive species.

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Term
Bulb
Definition
An organ used for vegetative reproduction, to ensure that perennial plants survive winter and reshoot in the next year. A short, flattened, or disk-shaped underground stem, with many fleshy scale-like leaves filled with stored food. Serves as a nutrient reserve for the meristem. Can have mini "bulblets" which fall off the bulb.
Term
Bulbous buttercup
Definition

Ranunculus bulbosus

Family Ranunculaceae. Has vegetative propagation with corms. Found in Ontario and Europe.

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Term
Bull thistle
Definition

Cirsium vulgare

Family Asteraceae. A perennial. Has a taproot.

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Term
Callaloo
Definition
A dish which incorporates the leaves of plants which may be considered weeds.
Term
Canada thistle
Definition

Cirsium arvense

Family Asteraceae. A creeping perennial. A weed found in North America and Australia which was introduced from Europe. Moved from France to Quebec, then spread across the continent to California. Has vegetative propagation with horizontal roots, which can go very deep.

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Term
Canola
Definition

Brassica rapa

Family Brassicaceae. As a volunteer, it is the second most common weed in Western Canada.

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Term
Carbon dioxide (CO2)
Definition
A resource that plants require, however they do not compete for it because it is not a limiting resource. It is in great quantity in the atmosphere, and more is made all the time.
Term
Caribou
Definition
In BC, populations were decreasing, so they brought more moose into the area. The moose do not compete with the caribou, but they are the preferred prey of wolves.
Term
Carotenoids
Definition
A light sensitive molecule in plants. Reflect yellow and red light.
Term
Caryophyllaceae
Definition
A family of plants. The pink family. Includes common chickweed, carnations, and white cockle.
Term
Chenopodiaceae
Definition

"Goose foot"

A family of plants. Includes common lamb's quarters, fig-leaved goosefoot, spreading atriplex, kochia, sugarbeet, quinoa, beets, and spinach. Leaves can resemble goose feet. Evolved to live in saline environments, such as sea coasts, or areas with little irrigation. Salts are pumped into the trichomes, to exude it from cells.

Term
Chlorophyll
Definition
A light sensitive molecule in plants. Reflects green light.
Term
Clarence Swanton
Definition
A friendly, passive guy. But he will go on a rant about how allelopathy is not as important a factor in interference as competition.
Term
Climax
Definition

Climax community

Stability

The seventh and final stage of the succession process. There is a stable climax community, which is usually a wooded forest. It can take 75 - 100 years to reach climax.

Term
Cocklebur
Definition

Xanthium sp.

Family Asteraceae. Has hooks on the seeds to aid with dispersal, by sticking to animals' fur and peoples' clothes.

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Term
Coleoptile
Definition
A protective membrane over the cotyledon of a monocot seedling. Helps it get through the soil.
Term
Coltsfoot
Definition

Tussilago farfara

Family Asteraceae. A creeping perennial. Has vegetative propagation with rhizomes.

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Term
Common burdock
Definition

Arctium minus

Family Asteracea. Has hooks on the seed to aid with dispersal, by sticking to animals' fur and peoples' clothes. The hooks were the inspiration for the invention of Velcro.

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Term
Common chickweed
Definition

Stellaria media

Family Caryophyllaceae. A winter and spring annual. Common in turf and winter crops. Has vegetative propagation with rhizomes. It looks fragile, but can be a persistent weed. Leaves are ovate, with a pointed tip, arranged oppositely. Prostrate growth habit, forming mats. Flowers are small. Petals are bifid, with 5 petals appearing like 10. Stem is hairy on one side only, with sides alternating by node. Can appear similar to scarlet pimpernel.

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Term
Common lamb's quarters
Definition

Chenopodium album

Fat hen

Family Chenopodiaceae. Closely related to quinoa and spinach. Name comes from when it grew in lamb's pastures. The tenth worst weed worldwide. Very common in all crops, anywhere soil is disturbed. A pioneer species; exploits resources quickly. Its WUE is 658; this poor water efficiency wastes water from the available pool, thus reducing water availability for neighboring plants. Has luxury consumption of nitrogen. Edible, and tastes like spinach. Has an erect growth habit. Produces salt bubbles on its surface, giving it a white sheen with a mealy texture. Leaves are alternating. Has small, green flowers. Seeds can survive for 1,700 years in the seedbank. Seeds must be near the surface to germinate. Can appear similar to fig-leaved goosefoot and spreading atriplex. 

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Term
Common mallow
Definition

Malva neglecta

Family Malvaceae. A perennial. Can have a deep taproot. If it has the space to, it will take on a bushy growth habit. If it is mowed down or cut, it grows in a prostrate growth habit. Leaves are kidney-shaped with irregular, jagged indentations. Flowers are small, with five petals, white with pink stripes. Some ornamental varieties have bigger flowers.

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Term
Common milkweed
Definition

Asclepias syriaca

Family Apocynaceae. Has vegetative propagation with rhizomes, which can reach 2 feet deep. Can produce latex and silk. Seeds can survive for 3 years in the seedbank. Has feathery appendages on the seeds to aid with dispersal on the wind. It was removed from the list of noxious weeds because it is a food source for the Monarch butterfly.

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Term
Common mullein
Definition

Verbascum thapsus

Devil's tobacco

Family Scrophulariaceae. A biennial weed, capable of very early start in the spring. It can grow back after a fire. Highly competitive. An early successional species. Can grow to be very tall. Has fuzzy leaves.

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Term
Common yellow woodsorrel
Definition

Oxalis stricta

Family Oxalidaceae. It is edible, and has a sour taste. It contains oxalic acid, which is why it is bad to eat too much of it. Leaves are trifoliate, with heart-shaped leaflets. Flowers occur singly, and have 5 petals. Sometimes leaves are mistakenly sold as clover shamrocks near St. Patrick's Day. Can appear similar to black medick.

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Term
Community
Definition
Different species assembled in a functional system. During the reaction phase of succession, one plant community is replaced by another. At the end of succession, there is a stable climax community.
Term
Competition
Definition

Indirect interference

The fourth stage of the succession process. A contributing factor of interference. The negative relationship between two species that is due to the exploitation of resources that are present in limited supply, and are needed for growth and development. Involves various species competing for resources and space. Some plants may grow faster than others. Competition occurs for three types of resources: water, nutrients, and light. The outcome of competition is decided by: weed density, weed distribution, spacing, time, and weed species. Competition for different types of resources is interrelated, because lack of one resource can diminish a plant's ability to acquire other resources.

Term
Conditionally dormant
Definition
When a seed can germinate under a narrow range of conditions.
Term
Conservation
Definition
Nature areas may be conserved for peoples' enjoyment. Weeds can interfere with these environments.
Term
Conventional tillage
Definition
Systems where soil is tilled on a regular basis. The succession process is continually reset. Pioneer species are found here chronically, and they enrichen the seedbank with their seeds over time. Tillage is often intended to kill weeds ("revenge tillage"), but it encourages them in the long run.
Term
Corm
Definition
The swollen base of a stem axis. Can be an organ for vegetative propagation. New corms form, and the old ones shrink and are brought underground. Similar to a bulb, but without layered leaves.
Term
Corn
Definition

Zea mays

Family Poaceae. A tall, dense crop. Early control of weeds allows the plants to produce enough foliage that they shade the soil, preventing further weed germination. Can experience nitrogen toxicity. Volunteer corn may be the main grass weed in Ontario, according to one Ridgetown professor. If it is artificially made to self-pollinate for multiple generations, the plants become inbred; they are short and low-yielding. Crossing two inbred lines produces a hybrid plant.

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Term
Cover crops
Definition
Plants grown in agricultural fields not for any harvested product, but for protection of soil and environmental conditions. Not a new idea, but is coming back into fashion recently. Sometimes weeds can have similar qualities to cover crops. Can become weeds themselves, especially multi-species cover crops. Increase earthworm populations. May trigger shade avoidance response in crops, from reflected light.
Term
Crabgrass
Definition

Digitaria sp.

Family Poaceae. An annual. Has vegetative propagation with rhizomes.

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Term
Cranberries
Definition

Vaccinium macrocarpon

Family Ericaceae. Grown in Muskoka, in flooded fields. Can experience aquatic weeds.

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Term
Creeping buttercup
Definition

Ranunculus repens

Family Ranunculaceae. A creeping perennial. Has vegetative propagation with stolons. Invades pastures. Can be toxic to horses and cattle. Forms a crown of trifoliate leaves. Can appear similar to tall buttercup, but it has more lateral branching. 

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Term
Creeping Charlie
Definition

Glechoma hederacea

Family Lamiaceae. A creeping perennial capable of nutrient foraging.

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Term
Creeping perennial
Definition

Wandering perennial

A perennial which has vegetative multiplication due to abscission or rotting. This is important even if there is no cultivation. It spreads out sideways. May have spontaneous dispersal by aerial bulblets, stolons, rhizomes, and roots. Many have the capacity for nutrient foraging.

Term
Critical period
Definition
The period of time during which crops must be weed-free to ensure yield is as high as possible. Occurs when the crop is still emerging, and has not yet formed a canopy. Control should be maximized at this time. After this period, weeds have less of an impact, because yield is already established.
Term
Cross-pollination
Definition
Sexual reproduction where pollen from one individual fertilizes an egg from another individual. It is common in most plants, but relatively rare in weeds. A pollen vector is required, such as wind, insects, or birds. Flowers may be showy to "advertise" to potential pollen vectors. Forces genetic recombination at each generation. Allows for greater gene exchange, and produces genetic diversity necessary for evolution. Isolated individuals cannot reproduce; a plant with obligate cross-pollination could never reproduce if it was the only plant in the area.
Term
Curled dock
Definition

Rumex crispus

Family Polygonaeae. A stationary perennial with a taproot. Can develop shoots from only the upper part of the taproot. Found near fences, in pastures, and no-till systems. Likes moist areas, such as near streams, but can grow elsewhere. Has long leaves with wavy edges and a leathery texture. There are red dots on the leaves. Has small greenish flowers. Fruits have floaters of trapped air, allowing them to float on water, and be carried by wind.

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Term
Dahlia
Definition

Family Asteraceae. Has vegetative propagation with corms.

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Term
Dandelion
Definition

Taraxacum sp.

Family Asteraceae. A stationary perennial. Has vegetative propagation with a vertical taproot. Can develop shoots from any part of the taproot when it is cut. Has a showy flower, despite reproducing mostly from apomixis. There are only a few races of dandelions, which are fixed clones. Has feathery appendages on the seeds (the pappus) to aid with dispersal on the wind, and may help seeds grip to animals. In one experiment, seeds were planted in various positions, and germination rates measured. The seeds germinated very poorly when planted upside-down; the pappus must have another function in ensuring the seed lands in the right orientation. 

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Term
Dicot
Definition

Broadleaf

A grouping of plants.

Term
Dispersal
Definition
Allows weeds to invade new sites. Many weed seeds are excellent travelers. Dispersal is done mostly by seeds, but it is possible for vegetative structures to be spread, contributing to dispersal. Prevents competition between seeds produced by the same mother plant. Includes natural and artificial dispersal, and vertical and horizontal dispersal.
Term
Disturbance
Definition

Nudation

The first stage of the succession process. Development of a bare site. Anything which kills vegetation. Can be a volcanic eruption, filling up of a lake, forest fire, tillage of a field, or use of herbicides.

Term
Dodder
Definition

Cuscuta sp.

Family Convolvulaceae. A parasitic plant. Almost like a disease. A noxious weed under the Weed Control Act. Not very common in Ontario. Grows near rivers. It is on the list of noxious weeds on the Ontario Seed Act. Can be a problem in alfalfa. It is more problematic in other countries, especially warm, dry climates. It sucks moisture from the host plant.

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Term
Dolly
Definition
A sheep which was cloned. The clone died of cancer because its telomeres were shortened.
Term
Dormancy
Definition
When a seed cannot germinate. Prevents viable seeds from germinating when conditions are adequate. Allows seeds to stay in the soil for long periods of time and survive. A survival mechanism. Wild plants without seed dormancy have less chance of survival because all the seeds would germinate at once, and the whole population could be wiped out. Can be caused by genetic and environmental factors including physiological, physical, morphological, and combinations of the three. Dormancy can be broken in different ways, depending on its cause. Includes primary and secondary dormancy.
Term
Early successional species
Definition

Intermediate species

Species which are found during the middle of the succession process, after pioneer species. Includes grasses, shrubs, pines, oak, and hickory. In abandoned agricultural fields, there are grasses and bushes. Includes biennial and perennial species. Mostly herbaceous. Found in no-till systems, pastures, turf, and orchards, where there is reduced disturbance.

Term
Earthworms
Definition
Good for the soil. Populations increase with no-till and cover crops. One farmer said that the ideal number was 10 earthworms in one shovel-full of soil. In turf grass, earthworms are undesirable because their tunneling makes a mess with close mowing, so they are killed with pesticides, sometimes pesticides intended for use on other pests.
Term
Eastern black nightshade
Definition

Solanum ptychanthum

Family Solanaceae. A weed which has poisonous berries. Can affect humans, livestock, and pets.

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Term
Ecesis
Definition
The third stage of the succession process. Involves establishment and initial growth of vegetation.
Term
Emergent weeds
Definition
Weeds which grow in wet areas, such as drainage ditches. Use of herbicides is dangerous, because water can be contaminated. Includes Phragmites.
Term
Field bindweed
Definition

Convolvulus arvensis

Family Convolvulaceae. An early successional species. A creeping perennial. A noxious weed under the Weed Control Act. Has vegetative propagation with roots and rhizomes. It will return even after it is pulled out. It is able to climb on other plants, and position its leaves more favourably for competition for light. Can look similar to wild buckwheat, but has a more variable leaf shape.

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Term
Field violet
Definition

Viola arvensis

Family Violaceae. Fruits have forced dehiscence.

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Term
Fig-leaved goosefoot
Definition

Chenopodium ficifolium

Family Chenopodiaceae. Leaves resemble fig leaves; there are indentations on the sides. Other than leaf shape, it appears similar to common lamb's quarters.

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Term
Fire
Definition
Can break seed dormancy in some species. The heat and charring reduces the seed coat. Some species are specifically adapted to germinate only after a fire.
Term
Fluorescent light
Definition
The light from fluorescent light bulbs has more green wavelengths. When you take a photo indoors using a camera intended for outdoor use, colours in the image will have a dull appearance, and look greenish.
Term
Forced dehiscence
Definition
A form of natural dispersal where seeds are expelled violently away from the mother plant. Found in wild cucumber, wood sorrel, and field violet.
Term
Forestry
Definition
When trees are cut down, weeds appear because the canopy is opened and they can grow quickly.
Term
Foxtail
Definition

Alopecurus sp.

Family Poaceae. The third most common weed in Western Canada.

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Term
Foxtail barley
Definition

Hordeum jubatum

Family Poaceae. A stationary perennial with shallow subterranean stems. Grows in clumps.

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Term
Germination
Definition
A critical stage in the life of a plant. It is the most vulnerable to environmental conditions. Steps include: imbibition, embryo metabolism, elongation of the radicle, and elongation of the shoot. Seeds germinate only when conditions are optimal, to increase chances of survival. Sometimes seeds don't germinate because: the seed is too young, the embryo is underdeveloped, the seed coat is too hard, there are internal inhibitors, or the seed is dead (non-viable). Main factors affecting germination are: light, oxygen, temperature, moisture, and depth in the soil. Crops are bred to germinate when they are planted. Weeds rely on environmental triggers to germinate.
Term
Giant foxtail
Definition

Setaria faberi

Family Poaceae. The ligule is ciliated, and the leaf blade is hairy on one side.

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Term
Giant hogweed
Definition

Heracleum mantegazzianum

Family Apiaceae. A weed that can cause skin irritation by making the skin sensitive to UV rays.

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Term
Giant ragweed
Definition

Ambrosia trifida

Family Asteraceae. Can grow to be fairly tall. Produces allergenic pollen. One grad student helped grow a plant in the greenhouse. He gave it a large container and plenty of fertilizer. It grew all the way to the greenhouse roof! The pollen from the plant had to be scooped off the ground, there was so much of it.

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Term
Golf
Definition
Aesthetic is important with golf courses. People involved in turf for golf are obsessed with making the lawns pristine.
Term
Green pigweed
Definition

Amaranthus powellii

Powell amaranth

Family Amaranthaceae. Stem is reddish at the top, with few hairs. There are tepals in the flower. Can appear similar to redroot pigweed, but the sheen on the leaf is shinier, though this can depend on soil nutrition. The inflorescence is less compact than in redroot pigweed, and more finger-shaped.

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Term
Hay fever
Definition
Allergy to pollen of weeds, including ragweed and some other species.
Term
Herbicides
Definition
Not as strong of a disturbance as tillage.
Term
Horizontal dispersal
Definition

Geographical dispersal

When seeds travel a short or long distance from the mother plant. Seeds may be carried by wind, water, or animals.

Term
Horsetail
Definition

Equisetum sp.

Scourbrush

Family Equisetaceae. A creeping perennial. Has vegetative propagation with rhizomes, forming a dense rhizome system that can compete with crops underground. Looks like sticks, with no leaves.

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Term
Imbibition
Definition
The first step in germination. When plant matter (usually a seed) absorbs water, and increases in volume. Used by ancient Egyptians to split rocks apart with wooden stakes. Allows plants to break apart cement. A dry seed regains moisture, and ends dormancy. The seed solubilizes its constituents, and begins chemical reactions necessary for germination. The seed coat breaks, and lets in oxygen for respiration. Seeds can produce a lot of power.
Term
Increased production costs
Definition
A negative aspects of weeds. Extra measures must be taken to control weeds, costing money.
Term
Interference
Definition

Competition

The negative relationship between two plants. The result of two phenomena: competition and allelopathy. It is the sum of these two factors. It is difficult to distinguish the contribution of each component to the overall outcome of interference. The effects of the two factors are difficult to separate experimentally.

Term
Animal power
Definition
Used to pull tillage implements. First recorded in 1,000 BC. Can use horses, cattle, or oxen.
Term
Canada Seeds Act
Definition
A law which ensures purity of seed for planting, and preventing of the spread of weeds. Regulates the seed industry. Enforced to avoid planting weed seeds along with crops. Includes tables of seed grade, based on total weed seeds in 500 grams of seed. Certified seed carries a blue tag, and has a lot less chance of having weed seeds than other seed.
Term
Canopy
Definition
Under a canopy, the R/FR is typically 0.5 - 0.75. Plants in these conditions have a higher Pr/Pfr ratio. The deeper in the canopy, the lower the value. Plants, such as weeds, living under the canopy suffer from two phenomena of shading: diminution of total light intensity, and alteration of light quality (reducing R/FR). Reduced R/FR light causes more Pr phytochrome, triggering multiple actions, including shade avoidance response. A canopy can be the best form of control, and may involve timing of cover crops. Weeds often appear in openings in the canopy.
Term
Common ragweed
Definition

Ambrosia artemisiifolia

Family Asteraceae. A noxious weed under the Weed Control Act.

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Term
Containment
Definition

Quarantine

The second phase of eradication. Ensuring there is no dispersal of the weed. Species disperse mostly by human means. For Striga in the USA, there was limitation on movement of soil, compost, crops, produce, and machinery such as implements, harvesters, tractors, and various tools.

Term
Control
Definition
An approach in weed management. Aims to reduce weed populations until it is not an economic problem. Generally involves medium- to low-intensity intervention. Time frame is not in the present, but in the short-term. Basically, a reactive management instead of proactive action. Uses mostly tillage and herbicides. Successful weed control is better achieved when more than one method is used; this is not always the case as the trend over the years has been to rely solely on herbicides. A continuous process; weeds don't go away in one year.
Term
Eradication
Definition
An approach in weed management. The word may mistakenly be used synonymously with "control". The total elimination of a weed species from an area, permanently. High intensity weed management over a prolonged time frame. Difficult, costly, and time-consuming to achieve. Involves getting rid of all vegetative plants, seeds, and other propagules, such as tubers. True eradication is realistic only for small-scale infestations in a limited size area. Striga was eradicated in the USA. Phases include survey, containment, and eradication per se.
Term
Eradication per se
Definition

Control of weeds

The third phase of eradication. For Striga in the USA, plants were eliminated as they were found, to prevent seed set. Herbicides were used in crops. Attempts at controlling the seedbank were made. Deaths in the seedbank was accelerated by triggering suicide germination without a host using ethylene, or accelerating mortality with the fumigant methyl bromide.

Term
Far red
Definition
The wavelength of far red light is around 705 - 740 nm. It is not absorbed for photosynthesis. It cannot be seen by humans. It causes Pfr to be converted into Pr.
Term
Goatgrass
Definition

Aegilops sp.

Family Poaceae. The seeds stick to the rachis and shatter. It is hard to clean its seeds from crop seeds.

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Term
History of weed management
Definition
As soon as agriculture started, weeds had to be managed. Their negative impact on crop growth and quality had to be reduced. When agriculture began, in 10,000 BC, the only method was hand pulling. Wood sticks and hoes were invented a few thousand years later. Use of animal power to pull tillage instruments was first recorded in 1,000 BC. In the 18th century, rotations with row crops developed in England, allowing inter-row cultivation, cleaning crops. Tillage implements pulled by tractors were first used in the 1920s, and use of chemical regulators and herbicides took off after WWII. Integrated weed management (IWM) is a recent concept of global weed control and crop management, being developed extensively. In the past 15 years, genetically engineered herbicide-resistant crops have caused a great transformation in global weed management.
Term
Interference with farm operations
Definition
A negative aspect of weeds. Too many weeds can slow mechanical harvest. Thorny weeds can slow manual harvest in orchards and vegetables.
Term
Jethro Tull
Definition

1674 - 1741

Lived in England, as a owner of a farm, who had people working for him. He traveled Europe, and became concerned that his farm was inefficient, so he invented the seed drill. It was unknown at the time whether weeds had any effect on crops, and Tull believed that they did. Jethro Tull is also the name of a band from the 1970s.

Term
Jewelweed
Definition

Impatiens capensis

Family Balsaminaceae. The seed pod builds up tension as it develops, then when cells disintegrate, suddenly seeds are flung away from the plant.

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Term
Kochia
Definition

Kochia scoparia

Tumbleweed

Family Chenopodiaceae. A bad weed in Western Canada and USA. Less common in Ontario, where its seeds may be found in alfalfa produced in Western Canada. The seeds are similar in size and shape to those of alfalfa, and can be difficult to clean out. Found on roadsides in Mexico and Arizona. Has resistance to many herbicides. Was once used as an ornamental. Seedlings have hairy leaves that grow in a rosette. Old and young leaves appear the same: narrow, with a smooth edge. The stem is ribbed, like in beets. Has a bushy and branchy growth habit. The plant forms into a ball shape, and as it matures, the base of the stem breaks. The plant rolls around, dispersing seeds as it tumbles. Sometimes tracks of seedlings can be seen, where a kochia plant has blown by.

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Term
Lady's thumb
Definition

Persicaria maculosa

Family Polygonaceae. An annual. One of the most common weeds worldwide. Found right after disturbances. There is a triangular or circular print of darker green on the leaf, like a "lady's thumb"; the spot may have to do with heat dispersal. Ochrea are smooth and hairless, with a few hairs at the tip. Not an elegant-looking plant; has a loose growth habit.

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Term
Late successional species
Definition
Species which are found late in the succession process, after early succesional species, near stability. Long-lived, slow-growth species, including trees. Includes mature oak and hickory forests. Develops after 30 - 60 years.
Term
Legumes
Definition
Tend to be good pasture plants. Includes many edible crops, and many weeds, including black medick.
Term
Lettuce
Definition

Lactuca sativa

Family Asteraceae. A delicate crop. It is slow-growing, and has a small root zone. Not a good competitor.

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Term
Lifecycle
Definition
A way to categorize plants, and weeds. Includes annual, biennial, and perennial plants. Sometimes the distinction is not clear, and can depend on the environment.
Term
Light
Definition
A factor in germination, and a resource for which plants can have competition. Germination can be influenced by light in different ways; it may stimulate it in some species, but inhibit or have no effect in others. Reaction to light in a seed is photosensitivity. Competition for light occurs at the canopy level, when the foliage of a plant is developed enough to shade the foliage of other plants. A reduction in the amount of light on a leaf results in chlorosis (yellowing), etiolation, and a reduction in the accumulation of dry matter. Plants that establish a positional advantage early in the season are more able to compete for light. Weeds that climb on other plants are able to position leaves favourably and intercept more light. Fortunately, weeds are not the only efficient competitors for light. Tall and dense crops such as corn and cereals can successfully compete for light with small plants, especially if established early. Early control of weeds in these crops allows them to produce enough foliage that shading of the soil will prevent further weed germination.
Term
Loose silky-bent
Definition

Apera spica-venti

Family Poaceae. A weed that was found in Norfolk County in 1976. Believed to have been introduced with rye brought from Europe.

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Term
Luxury consumption
Definition
When a plant can take up more of a nutrient than it needs for its growth. A number of weeds are capable of luxury consumption of nitrogen. The plant has a higher N concentration than it requires, and grows very quickly. Plants with luxury consumption probably do have a limit where nutrients become toxic, but it is a very high level, where it would probably cause soil toxicity. Example: in a sugar beet field infested with lamb's quarters (which has luxury consumption of N), increasing N fertilizer rates from 75 kg/ha to 150 kg/ha doubles the biomass of lamb's quarters, but has no effect on the sugar beets.
Term
Malvaceae
Definition
A family of plants. The mallow family. Includes common mallow.
Term
Microdisturbance
Definition
A small disturbance, such as an animal's hole or a single dead tree. Pioneer species can take advantage of them.
Term
Migration
Definition
The second stage of the succession process. Arrival of propagules.
Term
Mistletoe
Definition

Order Santales. A parasitic plant.

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Term
Moisture
Definition
A factor in germination. The seed must be imbibed with water. Seeds need a minimum of 14% water content before germination proceeds. Water in the seed is essential for biochemical processes. Excess soil moisture can kill seeds, and causing damping off. Algae on the soil surface can be a sign of excess moisture.
Term
Monocot
Definition
A grouping of plants. Includes grasses.
Term
Morphological
Definition
A factor in dormancy. The seed is shed, but the embryo is not yet fully developed. Development continues as the seed is in the soil. Only time can break the dormancy.
Term
Mowing
Definition
A type of disturbance. Low intensity disturbance.
Term
Mung bean
Definition

Vigna radiata

Family Fabaceae. Well documented as having a high swelling rate during seed imbibition. Some seeds were able to lift a beaker of sand while imbibing water.

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Term
Mustard
Definition

Sinapis sp.

Family Brassicaceae. A weed found in North America and Australia which was introduced from Europe. Moved from France to Quebec, then across the continent to California. A pioneer species. Cotyledons are kidney-shaped. Leaves at the bottom of the plant are longer and rounder than leaves near the tip of the plant, which are pointed. One plant germinated in July, and then flowered with only two leaves; has high plasticity.

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Term
Natural dispersal
Definition
Natural means of seeds dispersal. Special features on the seed help. Can include hooks on the seed (stick to animal fur and feathers), feathery appendages which act like a parachute (allow wind dispersal), ability to float on water (dissemination on irrigation water), and a thick seed coat (allows for survival of animal digestive systems, spreading in non-composted manure). Dispersal can be helped by the fruit. Includes forced dehiscence.
Term
Negative photosensitivity
Definition
When germination is inhibited  by the presence of light.
Term
Neutral photosensitivity
Definition
When germination is not affected by the presence or absence of light. Most crops have neutral photosensitivity.
Term
Nitrogen (N)
Definition
The nutrient for which there is the most competition. It is more mobile than other elements, and is easily taken by the roots of "greedy" weeds. Increasing N fertilizer rates is seldom helpful, because a number of weeds are capable of luxury N consumption. Some species can be suppressed by increased rates of N, such as prostrate knotweed. N fertilizer is one of the main costs of agriculture, used because plants respond to it well.
Term
No-till
Definition
Only a small furrow is opened for seed planting. A good thing which reduces disturbance of the land. There is less weed control, but improved soil properties. There are less weed propagules in the soil. Instead of burying weed seeds, they remain at the surface, where they are exposed to fluctuating temperatures, and have reduced viability. Eventually early successional species are found rather than pioneer species, making weeds harder to control. Reduces soil compaction and erosion, and increases organic matter. There is more stability in the system. An alternative to tillage is deep-rooted cover crops. A kind of "religion" to some people, who are highly opposed to tillage. In Australia it is very popular, and it can be hard to find a plow! In Ontario, it is adopted to a lesser degree.
Term
Non-dormant
Definition
When a seed can germinate under a wide range of conditions.
Term
Nutrient foraging
Definition
The soil is heterogeneous, with nutrient-poor and nutrient-rich pockets. When plantlets of creeping perennials reach areas of soil that are nutrient-rich, and they send nutrients back to other parts of the plant which are in nutrient-poor soil, through rhizomes or stolons. Roots have more branching in nutrient-rich areas.
Term
Nutrients
Definition
A resource for which plants can have competition. Nutrients for which weeds compete are those in limited supply, mostly nitrogen (N), phosphorus (P), and potassium (K). Most nutrient competition is for N, with competition for P and K less common. Competition occurs at the root level. Availability of nutrients can be affected by soil pH.
Term
Nutsedge
Definition

Cyperus sp.

Family Cyperaceae. A monocot, but not a grass. A creeping perennial. Has vegetative propagation with tubers. Can tower over lettuce. Can be a harmful competitor. Grown in some countries for its highly nutritious tubers, which may be called "nutlets".

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Term
Ochrea
Definition
A membrane over the node of plants in the family Polygonaceae. Wraps the axillary bud and leaf as it develops. Eventually expands and then degrades. May be absent on lower nodes, but will be seen on higher nodes.
Term
Onion
Definition

Allium cepa

Family Amaryllidaceae. Has vegetative propagation with bulbs.

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Term
Ox-eye daisy
Definition

Leucanthemum vulgare

Family Asteraceae. A dicot. A stationary perennial which uses a shallow, oblique subterranean perennating stem.

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Term
Oxalic acid
Definition
Acid which is used to clean or bleach decks. It is also found in common yellow woodsorrel.
Term
Ox
Definition
A castrated bull which was castrated as an adult. It is not a full castration.
Term
Oxygen (O2)
Definition
A factor in germination. Germination requires a lot of energy, from the oxidation of seed reserves via respiration. Germination cannot occur if there is not enough oxygen in the soil. Affected by soil porosity, depth of the seed, and density of soil microorganisms. O2 is consumed and CO2 is produced, which must be eliminated. If CO2 concentrations are too high, germination is inhibited. Amount of oxygen in the soil can be influenced by environmental conditions and agricultural practises. In flooded soil, or fields with green manure, oxygen can be reduced, inhibiting germination. Weeding increases soil oxygen, stimulating germination of seeds.
Term
Pappus
Definition
Papery tissue on the seed of Asteraceae plants. An energy investment which aids in dispersal.
Term
Parasitism
Definition
When one plant directly affects another by stealing nutrients. The parasitic plant attaches directly to the host plant, and accesses its photosynthates. Includes dodder, mistletoe, and Striga. The plant has no photosynthesis of its own, and may be yellow in colour.
Term
Perennial
Definition
A plant which grows back year after year. Lifecycle lasts more than two years. Perennial weeds require a different approach, and are often found in pastures and turf. Vary in ability to propagate themselves, depending on the type of vegetative organs they posses. Can have limited multiplication with offshoots, suckers, offsets, and ratoons. Includes stationary and creeping perennials.
Term
Pfr
Definition
The active form of phytochrome. Absorbs far red light. Upon absorption of far red photons, it is converted into Pr. Reacts to a lesser extent to red light. Pr is converted into Pfr upon absorption of red photons. In the dark, it is spontaneously, slowly converted into Pr at a linear rate. Triggers various physiological processes by altering gene expression.
Term
Photosensitivity
Definition
When a seed reacts to light. Includes positive, negative, and neutral photosensitivity. Reaction to light occurs only after the seed is imbibed. Crops are selected to have neutral photosensitivity. Germination of photosensitive species can be influenced by quality of light. Phytochrome controls germination based on R/FR ratio in the light, allowing it to sense if it is under a canopy, and the time of year. This prevents seeds from germinating in the wrong season, when other plants are well established, or when buried too deep in the soil. Light exposure can include even short bursts, such as during tillage.
Term
Phragmites
Definition

Family Poaceae. A tall grass which is an emergent weed, growing in wet ditches. It grows from the tropics to the arctic. Pulling it out makes no difference because it is a perennial. Herbicides such as RoundUP may be used on it, though there is risk of contaminating the water.

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Term
Physical
Definition
A factor in dormancy. The embryo is developed and there are no inhibitors in the seed, but the seed coat is hard, preventing water and oxygen from getting in. Dormancy can be broken by scarification which increases porosity of the seed coat.
Term
Physiological
Definition
A factor in dormancy. The embryo is fully developed, but there are chemicals in the seed which inhibit growth of the embryo. The compound is generally the plant hormone abscisic acid (ABA). Dormancy can be broken by alternating cold and warm temperatures over time, or stratification.
Term
Phytochrome
Definition
A family of light sensitive pigments that play a role in plant development and many other responses and bioprocesses. Enables plants to perceive the ratio of red light to far red light (R/FR). The ground, inactive state is Pr, and the active state is Pfr. It is synthesized in the Pr form. These two forms convert back and forth when they absorb red or far red light. A balance of the two forms forms based on the R/FR of light incident on the plant, as conversion reactions take place. Involved with plants being able to detect shading, neighboring plants, and day length. Confers seed photosensitivity. During the day, Pr and Pfr form an equilibrium based on perceived R/FR, but at night Pfr slowly converts to Pr by a natural spontaneous reaction with a linear rate. Under long days (short nights), there more increased Pfr in the morning. Under short days (long nights), there is reduced Pfr in the morning. This allows plants to perceive day length (and thus time of year). There are several types of phytochrome.
Term
Pigweed
Definition

Amaranthus sp.

Family Amaranthaceae. A pioneer species. Can grow very fast. Produces a canopy in 8 days, becoming a problem. Can appear similar to three-seeded mercury. One grower had it growing in strips in his field, as though it had been planted artificially; seeds had fallen off an uncleaned combine brought in from the USA. Some Jamaican species are grown as a crop, similar to amaranth.

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Term
Pineapple
Definition

Ananas comosus

Family Bromeliaceae. Has vegetative propagation with ratoons, which can form after harvest.

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Term
Pioneer species
Definition

Ruderal species

Species which are found early in the succession process, right after a disturbance. Includes annual plants, and perennial grasses and dicots. Generally annual, and short-lived. Adapted for sudden release of nutrients in the environment, such as after a fire. Exploit environment resources quickly. Have high plasticity. Can live in short-term openings in the land, such as flood plains. May appear every 3 - 5 years.

Term
Plasticity
Definition
Ability to change growth habit rapidly in response to stress. Weeds can be highly plastic; can produce seeds quickly if there is stress. This has lower productivity, but is safer.
Term
Poison ivy
Definition

Toxicodendron radicans

Family Anacardiaceae. A weed that can cause skin irritation. Leaf oils cause an allergic reaction on the skin, leading to itchy blisters. Takes 3 - 4 days for symptoms to develop. Scratching makes it worse. Grows as a vine. Leaves are trifolaite, variable in shape, with irregular indentations. The two leaflets to the sides may be asymmetrical, and the centre leaflet has a longer petiole.

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Term
Polygonaceae
Definition

"Many knees"

A family of plants. Has characteristically swollen nodes with ochrea. Includes curled dock, lady's thumb, prostrate knotweed, and wild buckwheat.

Term
Poinsettia
Definition

Euphorbia pulcherrima

Family Euphorbiaceae. It is very sensitive to daylength, requiring short days (long nights) to flower. In Ontario, to speed up the flowering process in time for Christmas, greenhouses use curtains to block out light. Even one flash of light can ruin the effect. Grows as a weed in New Zealand.

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Term
Positive photosensitivity
Definition
When germination is stimulated by the presence of light. Most weed species have positive photosensitivity. To prevent triggering germination of weed seeds, in Europe some farmers fave tried tilling at night; about half of studies found no effect.
Term
Potato
Definition

Solanum tuberosum

Family Solanaceae. Has vegetative propagation with tubers as well as rhizomes, which are called stolons. Daylength and biomass triggers growth of tubers. The eyes are the nodes on the tuber, with meristems that can sprout to form a new plant.

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Term
Pr
Definition
The ground state of phytochrome. The inactive form of the pigment. Absorbs red light. Upon absorption of red photons, it is rapidly converted into Pfr. Pfr is converted into Pr upon absorption of far red photons. In the dark, Pfr is converted into Pr in a slow, spontaneous reaction.
Term
Prevention
Definition
An approach in weed management. Stopping a weed problem before it becomes too much of a problem and invades a new location. Stopping weed dispersal. Generally involves low intensity measures. Action takes place in the present, but rewards are in the future, in the long-term. Can involve avoiding weed infested manure, compost, or straw, cleaning machinery between fields, buying certified seeds, cleaning seed, and killing weeds in ditches and hedgerows. Killing weeds in the natural environment might be harmful to wildlife. May be considered boring.
Term
Primary dormancy
Definition
When seeds are shed from the plant in a dormant state.
Term
Primary succession
Definition
The succession of plant species which occurs when a new environment forms where plants did not exist before. Includes a  lava field, a lake filling up, or glacial activity. Plants must come in from wind-blown seeds.
Term
Primitive tools
Definition
Used for weed management thousands of years ago. Weapons for hunting were repurposed to make early hoes.
Term
Primulaceae
Definition
A family of plants. The primrose family. Includes scarlet pimpernel.
Term
Propagules
Definition
Plant dispersal units. May be wind-blown. Can stay in the soil.
Term
Proso millet
Definition

Panicum miliaceum

Family Poaceae. A C4 weed. Its WUE is 267.

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Term
Prostrate knotweed
Definition

Polygonum aviculare

Family Polygonaceae. An annual. Found on turf and roadsides. Can tolerate stress, including a lot of trampling. Found in areas where there is a lot of foot traffic, such as on the edges of paths cutting through grass fields. Has a branchy, prostrate growth habit. Can have a more upright growth habit when found in cereal fields. Has a fibrous stem. Leaves are small and bluish. Small flowers form in axils, white, sometimes pink. It is less competitive with higher rates of nitrogen fertilizers.

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Term
Pulling weeds
Definition
The only method of weed management that existed for thousands of years, after the start of agriculture in 10,000 BC. Can be tedious work.
Term
Purple nutsedge
Definition

Cyperus rotundus

Family Cyperaceae. Has vegetative propagation with tubers. The tubers look like tiny potatoes, and can have up to six nodes.

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Term
Purslane
Definition

Portulaca oleracea

Family Portulacaceae. A C4 plant. It is edible, and eaten in salads and stew as a vegetable. High in minerals and vitamins. It is grown as a crop in the Mediterranean. May be used as an ornamental; wild plants have yellow flowers. It is adapted to warm, dry conditions. Can be a problem in strawberries, because it has late germination. Has a mostly prostrate growth habit, growing close to the soil. Can form thick mats. Has an extensive root system, in order to find water in moisture-limited conditions. The stem can sometimes appear red. Leaves arranged oppositely (two per node). Leaves are smooth and succulent; the plant uses them as a moisture reserve. It is a good system for studying shade avoidance response, because of its prostrate and branchy growth habit; you can watch where the plants will "crawl". It will grow away from green plastic objects, but not plastic objects of other colours.

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Term
Quackgrass
Definition

Elytrigia repens

Family Poaceae. A creeping perennial. Found in cooler areas around the world. Has vegetative propagation with rhizomes. Its rhizomes can reduce soil erosion. An aggressive weed. Releases toxic compounds from decaying dead material. Residues slow growth of alfalfa and clover, and reduce nodulation.

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Term
Racoons
Definition
Carry a parasite in their intestines, which is also found in their feces. If eaten by a bird or a frog, the parasite goes into the brain and causes problems. Another racoon may then eat the animal, usually head-first, thus getting the parasite.
Term
Ragweed
Definition

Ambrosia sp.

Family Asteraceae. Many people are allergic to its pollen. The economic threshold is one plant per square metre, which causes over 20% yield loss.

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Term
Reaction
Definition
The fifth stage of the succession process. The reaction to the replacement of plant species. Involves autogenic changes affecting the habitat, resulting in replacement of one plant community by another. Leads to development of a climax community in the stabilization phase.
Term
Red
Definition
The wavelength of red light is around 650 - 670 nm. It is absorbed for photosynthesis. It causes Pr to be converted into Pfr.
Term
Red/far red ratio (R/FR)
Definition

Light quality

The ratio of red photons to far red photons in light. Controls the levels of phytochrome types (Pr and Pfr), thus affecting various physiological processes in plants. Can affect leaf expansion, chloroplast development, stem elongation, root development, flowering, tillering, and apical dominance. In low R/FR light there is reduced lateral branching. Enables plants to distinguish between shade of a plant, and that of a non-plant object. Sunlight has high quality because it is red enriched with a high R/FR of about 1.2. When light passes through or is reflected off a plant canopy, some red light is absorbed for photosynthesis, but far red light is not used. Transmitted or reflected light has a lower quality, with a R/FR of 0.5 - 0.75. The low quality light which a  plant reflects is a function of the distance from the plant.

Term
Redroot pigweed
Definition

Amaranthus retroflexus

Family Amaranthaceae. Hairs are curled downwards. Leaves in seedlings have notches at the tips. Has diamond shaped leaves. Flowers are green, in a spiny inflorescence which hurts to touch. Similar-looking to other pigweeds. Can appear similar to nightshade. Its WUE is 305. This excellent water efficiency enables it to maintain growth even when water supply is becoming depleted.

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Term
Reduction of yield
Definition
A negative aspect of weeds. One of the first reasons why people want to get rid of weeds. Weeds compete with other plants, stealing sunlight, nutrients, and water. Reduction of yield is more intense depending on the number of weeds, type of weeds, and type of crop.
Term
Rhizome
Definition
An organ used for vegetative reproduction, to ensure that perennial plants survive winter and reshoot in the next year. An underground shoot. Has the internal structures of a shoot, not a root. Used for vegetative propagation by creeping perennials. Horizontal rhizomes develop from lateral buds on the shoot. They grow and can develop secondary rhizomes from lateral buds, and can also develop new shoots after it has turned upwards. In undisturbed soil, there is little development of new shoots from lateral buds, but if rhizomes are cut, such as from tillage, extensive growth from lateral buds can occur. Form tubers if the tip is enlarged. Usually white or yellow in colour, but turns green if exposed to light.
Term
Rice
Definition

Oryza sativa

Family Poaceae. There were some attempts to grow rice in flooded fields in Ontario, but it didn't work well. Flooded rice can experience aquatic weeds.

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Term
Roots
Definition
Because we cannot see them, we tend to underestimate their role. The root zones of neighboring plants compete for water and nutrients. An organ used for vegetative reproduction, to ensure that perennial plants survive winter and reshoot in the next year. Used for vegetative propagation by creeping perennials. Thickened roots develop after secondary growth in thickness of slender roots. Aerial shoots arise from buds on the roots, as well as additional thickened roots. Fragments of roots are regenerative, and the plant is encouraged by tillage. Includes vertical taproots, and horizontal roots.
Term
RoundUp Ready
Definition
The biggest development in weed management in the last 20 years. It has become conventional.
Term
Row width
Definition
A crop management factor which affects the outcome of weed management. Changes the distribution of plants, and affects canopy closure.
Term
Ryegrass
Definition

Lolium sp.

Family Poaceae. A turf and forage crop. Can be a competitive weed in wheat and other grains. Has cross-pollination. Seeds live for 4 years; if you can prevent recruitment of seeds for this amount of time, you can eradicate it. Grown in England and the Mediterranean. Important in Australia.

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Term
Scarification
Definition
Microbes and sand slowly abrading the seed coat until water and oxygen can get in. A way of breaking physically-caused dormancy in seeds. Porosity of the seed coat is increased. Can involve fire.
Term
Scarlet pimpernel
Definition

Agnalis arvensis

Family Primulaceae. The stem is not hairy. There are black dots on the leaf underside. Flowers are red with 5 petals. Appears similar to common chickweed, but has darker green leaves. Scarlet Pimpernel is also the name of a superhero from the 18th century.

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Term
Secondary dormancy
Definition
Sometimes seeds go through a period that will break dormancy, but not necessarily germinate at this time. If this is prolonged, such as by winter, then the seed goes back into a secondary dormancy. Prevents seeds from germinating at the wrong time of year. Can be caused by a cool, wet spring followed by a warm, dry summer.
Term
Secondary succession
Definition
The succession of plant species which occurs when all or most of the existing plants in an environment are killed. Includes forest fires, and tillage of fields. Plants can come from seeds in the seedbank, or from wind-blown seeds.
Term
Seed
Definition
A protected stage in plants; not as vulnerable as seedlings or grown plants. A portable form of genetic material. Protects genetic material from the environment. Physiologically inactive if dormant. Can survive for a long time: decades to centuries under certain conditions. Lifespan depends on species and environment. Seeds containing oils are subject to more oxidation than starch-filled seeds, and have shorter lifespans. Does not perform photosynthesis; it relies on stored reserves. Produced from sexual reproduction or apomixis. A better way of preserving genetic material than any form of vegetative reproduction: longer survival and better dispersal than vegetative parts. Numbers of seeds produced can be very high.
Term
Seed depth
Definition
A factor in germination. The larger the seed, the deeper it may be planted and successfully emerge. Has an effect on seed viability; the deeper the seeds are, the more viable they remain over time because they are less exposed to fluctuating temperatures. Tillage buries weed seeds, increasing their viability.
Term
Seed dormancy cycle
Definition
Seeds can cycle between dormant and non-dormant, having conditional dormancy between these. Environmental factors can enforce secondary dormancy.
Term
Seed drill
Definition
Invented by Jethro Tull. Allows seeds to be planted in rows, allowing for inter-row cultivation, cleaning the field of weeds. Before this invention, seeds were generally scattered in fields.
Term
Seed grade
Definition
Amount of weed seeds in grain reduces the grade of the seed.
Term
Seedbank
Definition
Dormant seeds present in the soil. Longevity is influenced by type of seed, weather patterns, tillage, predation, and death rate.
Term
Seedling
Definition
Young, small, vulnerable plants. Can be damaged by insects, animals, children, and other things.
Term
Self-pollination
Definition
Sexual reproduction where pollen from one individual fertilizes an egg from that same individual. Less common than cross-pollination in most plants, but is very common in weeds. Allows for reproduction of isolated individuals, such as pioneer plants in microdisturbances. There is no need for a pollen vector, or any showy flower parts. Flowers tend to be green and insignificant. There is some gene exchange between the two gametes, producing offspring with increased homozygosity compared to the parent plant. Offspring are not clones of the parent, but are very similar to it. Maintains stable genetic combinations within populations. Can cause inbred plants which express bad genes. Plants which normally self-pollinate do not experience this inbreeding depression.
Term
Sexual reproduction
Definition
Reproduction involving gametes. Flowers are complex, an investment of energy. There may be: pigment, nectar, smell, and showy parts. Always produces seeds. Produces increased diversity, a benefit compared to asexual reproduction. May be disadvantageous, because bad genes may be expressed. Includes self-pollination and cross-pollination. Most species have a mixture of self- and cross-pollination, which can be expressed as a percentage. Even a small percentage can mean a lot of plants, and can be significant for the process of evolution.
Term
Shade
Definition
The shadow of a plant. There is reduced photon flux (light intensity), as well as a decrease in R/FR (light quality). The difference in spectral composition is not visible to humans, but can be detected by plants. The light wavelengths are not as good for photosynthesis. Shade from other objects, such as buildings, or shade cloth, has a different wavelength composition than shade from plants. Plants can detect when they are in the shade using phytochrome, and respond with shade avoidance response.
Term
Shade avoidance response
Definition

Photomorphogenesis

Light affecting the shape of a plant. A response that a plant has to being in low R/FR light, such as when in the shade of a canopy. There are longer internodes, smaller leaves, and less roots. The plant is trying to become taller to avoid shading. Chloroplasts appear more yellow. Also occurs in plants which perceive light alteration coming from the sides, indicating that it is a response that also helps plants avoid competition. Two plants side by side, not shading each other, can cause shade avoidance response due to reflected light.

Term
Shade cloth
Definition
Black mesh which is propped over crops, to reduce the intensity, but not the quality, of light.
Term
Shallow subterranean stem
Definition
Used by some stationary perennials for propagatioon. The stem can be branched or unbranched, and there are adventitious roots. The primary taproot is not persistent. Annual regrowth occurs from buds on the perennating stem. Includes foxtail barley, broad-leaved plantain, and ox-eye daisy.
Term
Shedding seeds
Definition
Found in wild plants. Seeds are dispersed after they are produced. In crops, this is not found; seeds tend to stay on the head.
Term
Shepherd's purse
Definition

Capsella bursa-pastoris

Family Brassicaceae. A biennial. Early leaves are spoon-shaped, but then leaves that form later are a completely different shape, with many deep lobes. Texture of leaves is coarse and sandpapery, because the hairs are 5-pointed, branched like small stars; they are small and hard to see. Seed pods have a triangular shape with a notch at the tip. The name comes from when shepherds used purses made from the scrotums of rams, which resemble the triangular seed pods. Has small seeds, which struggle to emerge when planted 2 cm deep.

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Term
Shoots
Definition
An organ used for vegetative reproduction, to ensure that perennial plants survive winter and reshoot in the next year. Short stems with compact internodes at or just below the soil surface. Includes underground shots without internodes.
Term
Silverweed
Definition

Argentina anserina

Family Rosaceae. A creeping perennial. Has vegetative propagation with stolons.

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Term
Soil pH
Definition
Can affect the availability of nutrients.
Term
Soil type
Definition
A factor which affects the outcome of weed management. Affects the type of weeds, cooling, and the effect of some herbicides. Includes loam, clay, and muck soils.
Term
Soybean
Definition

Glycine max

Family Fabaceae. A self-pollinating plant. All individuals in a variety are genetically very similar. Flowering is triggered by daylength. Varieties are each adapted to different latitudinal daylength zones. One field was all matured and yellow, except for 4 - 5 green plants which were off-type varieties, not triggered to flower in that zone. Was not popular in Manitoba until recently, when varieties were developed that were adapted to that area.

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Term
Space
Definition
May be considered a resource for which plants must compete. Plants obtain resources (water, nutrients, light) by occupying space in the soil or air, so all competition may be considered competition for space.
Term
Spreading atriplex
Definition

Atriplex patula

Family Chenopodiaceae. Leaves are arranged oppositely, with variable shape. Can be difficult to identify. Appears similar to common lamb's quarters, but has a more branching growth habit, and a more pointed leaf.

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Term
Squirting cucumber
Definition

Ecballium elaterium

Family Cucurbitaceae. Seeds are sprayed out of the fruit with juice as it dehisces from the plant.

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Term
St. John's wort
Definition

Hypericum perforatum

Family Hypericeae. A perennial. An invasive species, introduced from Europe as an ornamental. Invades pastures and rangeland in the USA. Has a shrubby growth habit, with a stiff, woody stem. Leaves arranged oppositely (two per node). Leaves have tiny "perforations" where cells are translucent, appearing like holes; this is the reason for the species name. Flowers have many stamens. Contains toxic chemicals which can cause the skin to become photosensitive. Cattle and horses which eat it have weight loss nearly to death, as well as open skin sores. For humans it has medicinal properties as an anti-cancer drug and an antidepressant. Some farmers in BC grow and sell it for its medicinal values.

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Term
Stabilization
Definition
The sixth stage of the succession process. The reaction phase leads to development of a climax community.
Term
Stationary perenial
Definition
A perennial which is confined to the place where it germinated. Dispersion using vegetative means is only for short distances where the soil is disturbed due to cultivation. Either uses a shallow subterranean stem or a taproot.
Term
Stratification
Definition
Cold temperatures under moist conditions. A way of breaking physiologically-caused dormancy in seeds. Can cause cracks in the seed coat.
Term
Strawberry
Definition

Fragaria x ananassa

Family Rosaceae. A creeping perennial. Has vegetative propagation with stolons. Stolons can be a hindrance to production, growing between rows, and may be cut off. In the first year, there are annual weeds, and in the second year you get perennials.

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Term
Striga
Definition

Family Orobanchaceae. A parasitic plant. Causes enormous yield loss in corn, sorghum, millet, and other grass crops. Seeds can live in the seedbank for 10 years. Originated from Africa and Asia. Identified in 1955 in North Carolina, and in view of the enormous damage potential, a federally based eradication program was started in 1957. The program was deemed a success, despite having slow progress. In 1957, there were 450,000 infested acres. By 2007, there was a 99% reduction to less than 3,400 infested acres.

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Term
Succession
Definition
The normal transition over time of the plant species present in an ecosystem, following disturbance of the land. The change in species over time is prescribed, and tends to follow a script. Begins with a disturbance, and ends with climax or stability. Plants go through pioneer species, early successional species, and late successional species. Stages include: disturbance, migration, ecesis, competition, reaction, stabilization, and climax. Types of plants in the succession depends on the region, soil type, and climate. Important in ecology, with implications in weed science. By cutting trees and opening land for agriculture, the ecosystem is disrupted and succession begins. Includes primary and secondary succession.
Term
Sulfuric acid
Definition
It was once sprayed onto cereals to kill weeds. Broadleaf weeds would die, but the cereals are only burned and still able to recover. Has a bad effect on soil pH and other soil factors.
Term
Sunflower
Definition

Helianthus sp.

Family Asteraceae. The fruit is the kernel, which lacks a pappus.

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Term
Sunlight
Definition
In direct sunlight, the R/FR is typically 1.2. Plants in these conditions have roughly equal amounts of Pr and Pfr.
Term
Survey
Definition
The first phase of eradication. Scouting and mapping. For Striga in the USA, it was done in 36 counties. Landowners cooperated by checking land. Scouts were sent on foot and horseback. Rewards were offered to those who reported it. There was a need to be precise, taking care not to miss an area.
Term
Tall buttercup
Definition

Ranunculus acris

Family Ranunculaceae. Invades pastures. Grows into one tall stem with branches at the top. Leaf is unifoliate, but deeply lobed. Can be toxic to horses and cattle. Can appear similar to creeping buttercup, but has less lateral branching. Petals overlap more so than in creeping buttercup, but this is an unreliable distinguisher.

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Term
Taproot
Definition
Used by some stationary perennials for propagation. Can be more or less branched. In undisturbed soil, aerial shoots develop from buds near the soil surface. Some species can develop shoots from any part of the taproot after it is cut, others only from the upper part of the taproot.
Term
Technology
Definition
Tractors now have GPS, and can be self-driving. There has been development of tiny robots which pollinate flowers.
Term
Temperature
Definition
A factor in germination. For each species, there is a temperature at which germination is optimized, as well as an upper and lower limit outside of which germination cannot occur. Soil temperature will condition the pattern of germination of certain species. Weeds that germinate at lower temperatures will appear earlier in the season. Other species require warmer soil temperature to germinate, and may appear after the first flush of weeds has been eliminated, making season-long treatment necessary.
Term
Tendril
Definition
A long, slender appendage that can react to contact, and wrap around objects, helping plants climb. Modified leaves. Found in tufted vetch.
Term
Terrestrial
Definition
Most plants are terrestrial and grow on land.
Term
Three-seeded mercury
Definition

Acalypha rhomboidea

Copperleaf

Family Euphorbiaceae. Cotyledons are obicular. First leaves have indentations which are forward-pointing. Flowers form clusters in the axils. Grows on the edges of shaded, moist areas, such as forests. Has natural tolerance to Pursuit, and can be found in agricultural fields. Can appear similar to pigweed.

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Term
Threshold
Definition
The level of a pest where it is economic to control it. Established scientifically. Depends on type of pest, and the treatment. Weeds vary so much that it is difficult to produce guidelines.
Term
Time
Definition
A factor which affects the outcome of competition. Includes duration of competition, and when it begins. The earlier weeds emerge, and the longer they are present, the higher the yield reduction. Timing relative to the crop is one of the most important predictors of yield loss. Weeds that emerge with the crop have more impact: this is the critical period. Weeds with delayed emergence give advantage to the crop.
Term
Tree planting
Definition
After an area is deforested, the lumber company may hire youths to replant trees.
Term
Trees
Definition
Slow growing and slow to establish. Have high competitivity because they produce a canopy, and have a large root zone.
Term
Tuber
Definition
An organ used for vegetative reproduction, to ensure that perennial plants survive winter and reshoot in the next year. The swollen part at the end of a rhizome.
Term
Tufted vetch
Definition

Vicia cracca

Family Fabaceae.  A legume. A perennial with rhizomes. Flowers are purple and symmetrical, arranged all facing one direction on the inflorescence. Can climb on other plants for support, using its tendrils, however it is not a parasitic plant. Often found near fences, in no-till systems, and pastures. It was introduced as a forage crop, but was replaced by more productive, higher yielding plants. Afterwards, it persisted as a weed. Has good qualities as feed for livestock. It is naturally tolerant of RoundUp, however it can be controlled by some other herbicides. Can be a problem in no-till soybeans.

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Term
Vegetative reproduction
Definition
Reproduction using vegetative organs. Asexual reproduction, without pollen. There is no exchange of genetic material, and offspring are clones of the parent plant. Mostly prevalent in perennial plants, but found in some annual species. Perennial species use specialized organs for vegetative reproduction, ensuring the plant will survive winter and reshoot in the next year: rhizomes, tubers, stolons, bulbs, roots, and shoots. The organs must contain specialized buds from which new growth will be initiated, and enough food reserves to ensure winter survival. The main advantage is an early start in the spring. Ensures consistency, and rapid establishment and recovery from control. Less complex than sexual reproduction; no need for elaborate flowers and specialized organs. Perennials often have the capacity to grow rapidly after control measures that cause defoliation, such as mowing, fire, cutting, or herbivory. The plant re-emerges from protected buds. Rhizomes and stolons may give creeping perennials the capacity for nutrient foraging. Compared to seeds, it is a poor way to preserve genetic material.
Term
Velvetleaf
Definition

Cissampelos pariera

Family Menispermaceae. A pioneer species. Can grow to be very tall, and may be a problem in corn. A highly competitive species. Seeds can survive for 40 years in the seedbank.

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Term
Vertical dispersal
Definition
When seeds go into the soil seedbank. Dispersal through time. Seeds do not travel far from the mother plant.
Term
Viability
Definition
How long seeds can survive in dormancy. Species, coat constitution, maternal effects, and environmental conditions can have an influence. Seeds with higher oil content have less capacity for long life than those with starchy content, due to oxidation. Seed depth has a positive effect; tillage increases viability of weed seeds by burying them.
Term
Volunteer plants
Definition
During harvest, some seeds fall on the ground, and sprout in the next year. Crop plants which grow outside of their season. Considered weeds. Can cause problems with the crop. Includes volunteer potato, soybeans, cereals, corn, and rye.
Term
Water
Definition
A resource for which plants can have competition. It may be limiting in supply when rainfall is limited. Irrigation of crops ensures that water availability to plants is less variable, but does not remove competition for water, because weeds may still access the water. Availability of water to plants is governed by three mechanisms: natural rainfall, placement of roots and their growth rate, and water utilization efficiency of each species. Competition occurs at the root level, and starts as soon as the roots of one plant invades the root zone of another. Competition happens well before foliage is highly developed. Water competition tends to occur in the summer, and is more of an issue in arid places or during droughts. When water is limited in supply, weeds can further reduce water available to the crops, to a level which affects yield. When plants do not get enough water, they wilt, reducing their ability to absorb nutrients and light. Some plants can sustain photosynthesis for longer in drought conditions than others, and have an advantage for competition.
Term
Water utilization efficiency (WUE)
Definition
The amount of water, in grams, a plant needs to produce one gram of biomass. A mechanism which governs water availability to plants. Plants vary in their rate of water conversion into biomass. Some species require much more water than others to produce the same biomass. Plants that produce more biomass per unit of water are considered more efficient. C4 species are more efficient than C3 species. Species with very different WUE values can still compete, using different strategies.
Term
Waterhemp
Definition

Amaranthus tuberculatus

Family Amaranthaceae. A successful weed species with cross-pollination. It is not tall enough to compete with corn. An invasive species. Found in USA, and starting to be seen in Ontario. One grower in Quebec found seeds stuck in a used combine brought from USA.

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Term
Waterways
Definition
There is a system of bureaucracy that prevents pollution of waterways in Ontario, including use of herbicides near streams. It was difficult for one group to have a trail officially connected along the Speed River due to red tape.
Term
Weed
Definition
A plant that is not wanted because it interferes with human activities. Humans decide whether a plant is a weed or not; by definition it is anthropomorphic. Some species are considered weeds in some situations, but not in others. Any species can be a weed, however we tend to notice that some species come back more often than others. Those species have characteristics that help them invade crops or other locations. They are opportunistic, and unslightly. Their effect is a question of their context. The study of the biology of weeds helps us understand how to manage them. Negative aspects include: reduction of yield, increase in production costs, reduction of quality of products including contamination, interference with harvest operations, harbouring pests and diseases, allergies and toxins to humans and animals, obstruction of agricultural drains, obstruction of driver vision at intersections, aesthetics, and social concerns. There are some benefits to weeds. Humans have struggled with weeds since the beginning of agriculture. Weeds are adapted to disturbed environments, appearing at the beginning of the succession process, when humans alter plants in an area for their own activities. Grow spontaneously in crops. There are weeds because nature wants to go back to a stable stage of succession. Nature pulls in one direction while farming pulls in another. Not necessarily native plants; most are introduced. In North America, 39% are native, 3% from Central and South America, 35% from Europe, and the remaining 23% are found worldwide. Weeds have moved a long way, and are still moving. Most are flowering plants. Many are closely related to crops. Includes parasitic plants. Account for 0.1% of total plant species; around 390 species. Many are sold as ornamentals or medicinal plants. May be classified by: dicot/monocot, lifecycle, and terrestrial/aquatic.
Term
Weed Control Act
Definition
Forces municipalities and landowners to control weeds deemed a threat to agricultural land. There is zero tolerance for prohibited noxious weeds. Currently there are 24 weeds on the list, with various reasons for being on the list. Includes field bindweed, dodder, and common ragweed. Each county and municipality has a weed inspector who investigates complaints, and may order you to clean up weeds, or cost charges to your taxes. There is also a provincial chief inspector. There are some tolerance levels for noxious weeds.
Term
Weed density
Definition
A factor which affects the outcome of competition. The number of weeds per unit area. Units include weeds/m2, or weeds/m of row. The higher the density, the higher the yield loss. This relationship is linear at first, then curves downwards, because weeds start to compete with each other. An action threshold can be developed using this metric. Weed biomass may be a better indicator of weed stress.
Term
Weed ID Guide for Ontario Crops
Definition
A book by Mike Cowbrough, edited by Francois Tardif. From University of Guelph. The professors thought that Ontario needed a better guide for weeds. They added some weeds, and removed some others, from an existing book.
Term
Weed management
Definition
Makes use of a variety of methods. Historically began when agriculture started in 10,000 BC. Incorporates concepts of biology. Factors affecting management include: environmental conditions (moisture), weed type, soil type, and crop management (row width). Depending on how and when methods are used, three approaches can be distinguished: prevention, eradication, and control.
Term
Weed species
Definition
A factor which affects the outcome of competition. Weeds vary in competitiveness due to morphology (tall versus short), and physiology (C4 vs C3). Characteristics of competitive plants includes: rapid canopy expansion, large leaves, mosaic leaf arrangement, climbing, high biomass allocation to the stem, rapid shade response, and sometimes, C4 physiology.
Term
Wheat
Definition

Triticum sp.

Family Poaceae. Can be planted in the fall (winter wheat) or spring (spring wheat). If the weather is wet before harvest, the seeds can germinate on the head. Its WUE is 557.

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Term
Wheat rust
Definition
A plant disease which can be harboured on grassy weeds.
Term
White clover
Definition

Trifolium repens

Family Fabaceae. A perennial.

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Term
White cockle
Definition

Silene latifolia

Family Caryophyllaceae. A perennial. Found in pastures and no-till systems. Leaves are opposite with a short petiole. The flower has a hairy bladder-like structure at the base, with protruding white petals.

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Term
Wild buckwheat
Definition

Fallopia convolvulus

Family Polygonaceae. An annual. It is quite common worldwide. The fourth most common weed in Western Canada. Appears when cereals begin to mature. The first leaves are heart shaped, pointed with two rear lobes. Can appear similar to field bindweed, but it has ochrea.

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Term
Wild carrot
Definition

Daucus carota

Family Apiaceae. The same species as carrot. Can harbour many of the same pests.

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Term
Wild cucumber
Definition

Marah sp. 

Family Cucurbitaceae. Fruits have forced dehiscence.

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Term
Wild garlic
Definition

Allium ursinum

Family Amaryllidaceae. Has vegetative propagation with bulbs. A creeping perennial. Produces small bulbils in the inflorescence.

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Term
Wild mustard
Definition

Sinapis arvensis

Family Brassicaceae. Closely related to canola and mustard. Cotyledons are kidney-shaped, like in canola seedlings. Leaves at the base are long and rounded, but near the tip they are more triangular.

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Term
Wild oats
Definition

Avena sp.

Family Poaceae. The most common weed in Western Canada. Has a large seed, and can emerge when planted as deep as 17.5 cm. Seeds have bristles and awns which react to moisture change by twisting around; this allows the seeds to roll, twist, and drill into the soil. The seeds spin if the awn gets wet. Cells on one side of the awn react to moisture differently than cells on the other side, producing the twisting motion. A single plant growing in the absence of competition can develop 86 km of roots in less than 3 months.

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Term
Wild parsnip
Definition

Pastinaca sativa

Family Apiaceae. A weed that can cause skin irritation.

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Term
Wilting
Definition
A plant's reaction to not acquiring enough water. The stoma of the leaves shut to reduce water loss, and photosynthesis stops. The plant will eventually dry out and die.
Term
Wood sorrel
Definition

Oxalis acetosella

Family Oxalidales. Fruits have forced dehiscence.

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Term
Y2K
Definition
In 1999, it was thought that the turn of the millennium would disrupt computers, and there was a whole industry for making computers "Y2K compatible". Nothing actually happened to computers in 2000.
Term
Yellow foxtail
Definition

Setaria pumila

Family Poaceae. Has silky hairs.

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Term
Yellow nutsedge
Definition

Cyperus esculentus

Family Cyperaceae. An aggressive perennial. Releases toxic compounds from decaying dead material.

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